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THE PHYSICAL EVOLUTION OF THE HUMAN SPECIES AND OF HUMAN RACES SECTION ONE

The teachings of science that relate to the origin and history of the human species constitute for us the most important part of the whole doctrine of organic evolution and now, having completely outlined this doctrine as a general one, we are brought to the point where we must deal frankly and squarely with the insistent questions arising on all sides as to the way that mankind is involved in the vast mechanism of nature's order.
These questions have been ignored heretofore, in order that the natural history of animals in general might be discussed without any interference on the part of purely human interest and concern. It now becomes our privilege, and our duty as well, to employ and apply the principles we have learned in order to understand more completely the origin of the human body as an organic type, the history of human races, the development of human faculty and of social institutions, and the evolution finally of even the highest elements of human life. These are scientific problems, and if we are to solve them we must employ the now familiar methods of science which only yield sure results.

We must not underestimate the many difficulties to be encountered, for the field before us is a vast territory of complex human life and of manifold human relations. Without prolonged exercise in scientific methods, it is impossible to view our own kind impersonally, as we do the creatures of lower nature. Furthermore it seems to many that an analysis of human life and biological history, even if it is possible, must alter or degrade mankind in some degree; this is no more true than that a knowledge of the principles of engineering according to which the Brooklyn Bridge has been constructed renders that structure any different or unsafe for travel. Man remains man, whether we are in utter ignorance of his mode of origin, or whether we know all about his ancestry and about the factors that have made him human.
It is because our species appears to occupy a superior and isolated position above the rest of nature that the mind seems reluctant to follow the guidance of science when it conducts its investigations into the history of seemingly privileged human nature. And it is feared also, that if evolution is proven for man as well as for all other kinds of animals, our cherished ideas and our outlook upon many departments of human life must be profoundly affected. This may be so, but science endeavors only to find out the truth; it cannot alter truth, nor does it seek to do so. We might well wish that the world were different in many respects and that we were free from the control of many natural laws besides that of evolution, but if the real is what it is, then our duty is plain before us; as we think more widely and deeply on the basis of ripened experience, it becomes ever clearer that a knowledge of human history gives the only sure guidance for human life.

To the zoölogist it seems strange that so many are opposed to a scientific inquiry into the facts of human evolution, and to the conclusions established by such an inquiry,--though, to be sure, this opposition is directly proportional to ignorance or misunderstanding of the nature and purpose of scientific investigation and of human evolution. The naturalist comes to view our species as a kind of animal, and as a single one of the hundreds of thousands of known forms of life; thus the question of human origin is but a small part of organic evolution, which is itself only an episode in the great sweep of cosmic evolution, endless in past time and in the future. Were we some other order of beings, and not men, human evolution would appear to us in its proper scientific proportions, namely, as a minute fraction of the whole progress of the world.

While the foregoing statements are true, it is nevertheless right that a close study should be made of the particular case of mankind. No doubt much of the naturalist's interest in nature at large is due to his conviction that the laws revealed by the organisms of a lower sphere must hold true for man, and may explain many things that cannot be so clearly discerned when only the highest type is the subject of investigation. It is only too evident that little more than a general outline can be given of the wide subject or group of subjects included under the head of human evolution.
We must divide the subject logically into parts, so that each one may be taken up without being complicated by questions relating to topics of another category, although the findings in any one department must surely be of importance for comparison with the results established in another section; for if evolution is universally true, the main conclusion in any case must assist the investigation of another, just as comparative anatomy and embryology supplement and corroborate each other in the larger survey of organic evolution. As before, the illustrations of each department of the subject must be selected from the stock of everyday observation and information that we already possess, for we gain much when we realize that evolution includes all the happenings of everyday life and thought, as well as the occurrences of the remote past.

For the present, then, the questions relating to the higher aspects of human life must be put aside, only that they may be taken up at the last. Social evolution likewise finds its place in a later section, after the phenomena of mind and mental evolution receive due attention and description. At the present juncture, the human species presents itself as a subject for organic analysis and classification, merely as a physical organism. Just as the study of locomotives must begin with the detailed structure of machines in the workshop before they can be profitably understood as working mechanisms, so the physical evolution of mankind must first be made intelligible before it is possible to prosecute successfully the studies dealing with the psychology, social relations, and higher conceptions that seem at first to be the exclusive properties of our species.

The problems of physical evolution of man and of men fall into two groups. Those of the first deal with the origin of the human species as a unit, and its comparative relation to lower organisms, while those of the second part are concerned with the further evolution of human races that have come to be different in certain details of structure since the human type as such arose. In the first part, all men will be assumed to be alike and the members of a homogeneous species whose fundamental attributes are to be compared with those of other animals; only afterwards will attention be directed to the differences, previously ignored, that divide human beings into well-marked varieties. It must be evident even at this point that the mode of evolution demonstrated by the first investigation will be likely to bear some close relation to the methods by which human races have evolved to their present diverse anatomical situations.

The foregoing classification of the problems concerned with the nature and origin of the human species renders it possible to restrict the immediate inquiry to a definite and precise question. It is this: does the evidence relating to the physical characteristics of our species prove that man is the product of a supernatural act of creation, or does it show that man's place in nature has been reached by a gradual process of natural evolution? In order to obtain an equally precise and definite answer to this question, referring to the particular case of most concern to us, it is obvious that the method to be employed is the one which has given us an understanding of organic evolution as an all-inclusive natural process. The data must be verified, related, and classified, so that their meaning may be concisely stated in the form of scientific principles. What are the facts of human structure, comparatively treated? How does the human body develop? Does palĉontology throw any light on the antiquity of man? Do the rules of nature's order control the lives of men? Our course is now clear; we shall take up serially the anatomy, embryology, and fossil history of the human species, in order to see that there is ample proof of the actual occurrence of evolution, and then, as before, we may look about for the causes which have produced this result by natural methods.

While it is necessary to treat the subject directly, namely, by examining the actual evidences relating to the particular case in question, it is worthwhile before doing so to point out that, as the whole includes a part, human evolution has already been proved beyond question. This conclusion must be accepted, unless reasons can be given for excluding mankind from the rest of the living world as an absolutely unique type, supreme and isolated because of some peculiar endowments not shared with the rest of animate nature. If these reasons are lacking, and the unity of organic nature be recognized, human evolution cannot be denied unless some interpretation more reasonable and logical than evolution can be given for the whole mass of facts exemplified and discussed in the foregoing chapters. We may accordingly approach the main questions by asking if there are any reasons for regarding the human species as a unique and isolated type of organism.

At the outset, we must recognize that in so far as the human body is material, its movements and mass relations are controlled by physical principles, like all other masses of matter. It is well, indeed, that this is so, for if gravitation and the laws of inertia were not consistent and reliable principles holding true at all times and not intermittently, it would be difficult to order our lives with confidence. In the next place, the general principles of biology hold true for the structure and physiology of the human species as they do for all other living things. A human body is composed of eight systems of organs, whose functions are identical with the eight vital tasks of every other animal. All these organs are made up of cells as ultimate vital units, and the materials of which human cells are composed belong to the class of substances called protoplasm. Human protoplasm, like all other living materials, must replenish itself, and respire and oxidize in obedience to biological laws that have been found to be uniform everywhere. Thus the human organism is no more unique in fundamental organic respects than it is apart from the world of physical processes and laws.

How does the matter stand when the general structural plan of a human being is examined? Is it entirely different from everything else? It is a fact of common knowledge that the human body is supported by a bony axis, the vertebral column, to which the skull is articulated and to which also the skeletal framework of the limbs is attached. These characteristics place man inevitably among the so-called vertebrata; he is certainly not an invertebrate, nor is the basic structure of his body such that a third group, outside the invertebrata and vertebrata, can be made to include only the single type--man.

Passing now to the classes that make up the group of vertebrates, we meet first the lampreys or cyclostomes without jaws, and the others with jaws, such as the fishes, amphibia, reptiles, birds, and mammals, each class distinguished by certain definite characters in addition to the vertebral column. The fishes have gills and scales; amphibia of to-day are scaleless, and they are provided with gills when they are young and lungs as adults; reptiles have scales and lungs; birds are warm-blooded and feathered; while mammals are warm-blooded and haired. Is the human species a unique kind of vertebrate, or does it find a place in one of these classes? The occurrence of hair, of a four-chambered heart which propels warm blood, of mammary glands, and of other systematic characters marks this species as a kind of mammal and not as a vertebrate in a section by itself.

The members of the class mammalia differ much among themselves; and now that we recognize clearly that man is a mammalian vertebrate, the next question is whether an order exists to which our type must be assigned, or whether we have at last reached a point where it is justifiable to establish an isolated division to contain the human species alone. We are familiar with many representatives of different mammalian orders and with the kind of structural characteristics that serve as convenient distinctions in denoting their relationships.
Horses and cattle, sheep, and goats and pigs resemble one another in many respects besides their hoofs, and they form one natural order; the well-developed gnawing teeth of rats and rabbits and squirrels place these forms together in the order rodentia; the structures adapting their possessors for a flesh-eating and predatory life unite the tribes of the lion, wolf, bear, and seal, in the order carnivora. Among these and other orders of mammalia is one to which the lemurs, monkeys, and apes are assigned, because all these forms agree in certain structural respects that place them apart from the other mammalia, in the same way, for example, that the races of white men may be recognized as a group distinct from the black and red races. But comparative studies, prosecuted not only by those who have been forced to adopt the evolutionary interpretation, but also by believers in special creation like Linnĉus and Cuvier and other more modern opponents of evolution, have shown that the peculiar qualities of this order are shared by the human species.
Indeed, the name of primates was given to this section by Linnĉus himself, because the human body found a place in the array which begins at the lower extreme with the lemurs and the monkeys and ends with man at the other end. Again it is found that no separate order of mammals exists to include only the genus _Homo_.

To one unacquainted with the facts of vertebrate comparative anatomy, the distinguishing characteristics of the primates seem to be trivial in nature. It is surprising to find how insignificant are the details to which appeal must be made in order to draw a line between our own division of mammalia and the others. It is well to review them as they are given in the standard text-books of comparative anatomy. Primates are eutheria, or true mammalia possessing a placental attachment of the young within the parent.
The first digits, namely, the "great toe" and the "thumb," are freely movable and opposable to the others, so that the limbs are prehensile and clasping structures; usually but not always the animals of this order are tree-dwellers in correlation with the grasping powers of the feet and hands. The permanent teeth succeed a shorter series of so-called "milk teeth," and they are diverse in structure, being incisors, canines, or "eye teeth," premolars, and molars; the particular numbers of each kind are almost invariable throughout the order and markedly different from those of other orders. The number of digits is always five, and with few exceptions they bear nails instead of claws. The clavicles, or "collar bones," are well developed in correlation with the prehensile nature of the fore limbs; a bony ring surrounds the orbit or eye socket. Finally there are two mammary glands by which the young are suckled. It is because any other details of difference between man and other forms are far less marked than the agreements in these respects, that the human species must be regarded as a primate mammalian vertebrate.